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I believe the rules that allow them to fly in formation are local rules (what is the bird in front of me doing?). I wonder if it is a local rule that allows them to do these proportions as well.

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I think more in terms of System 1 and System 2. Most bias is a system 1 issue although bias is an issue that influences more deliberative thought. The pigeons are obviously in system 1. They are just being reinforced for choosing to switch over time. Humans don't seem to learn from the reinforcement in the same way.

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